A while back, Chris over at Ediacaran put up a post in his “Paleoporn” series describing the environment and sedimentology of two sites, in the same formation, where the same Cambrian trilobite is found. His post is a description of the orientation of the molts in the two different environments. In the comments, we discussed the curious habit of finding inverted cephalons, ie, ventral up cephalons while the rest of the critter is found in its normal position(dorsal up).
In a thin bed of the Clays Ferry Formation in Anderson Co. Kentucky, I find numerous partial molts from the trilobite Isotelus gigas. Some of the bedding planes are almost entirely composed thoracic segments, cephalons and cranidiums, and pygidiums from this large asasphid. Occasionally, one comes across the curious preservation described above.
During ecdysis, the cephalic sutures of the trilobites rupture allowing the critter to escape from the front of its carapace. Sometimes during the egress, the critter will push the cephalon upside down. Hence, what we have here.
But, I brought home anyway. While hitting it with a scribe, I noticed the ventral, but I thought that it was trash and nearly blew through it… as more became apparent, I slowed up a bit(word to the wise).
More on that later.